Also see my google scholar page


  • Botero, C. A., L. J. Harmon, and Q. Atkinson. 2017. The promise and limits of eco-evolutionary studies of human culture. Religion, Brain & Behavior 7 (2), 153-155. link - pdf.

  • Boucher, F. C., V. Démery, E. Conti, L. J. Harmon, and J. C. Uyeda. In press. A General Model For Estimating Macroevolutionary Landscapes. Systematic Biology. link.

  • Caetano, D. S. and L. J. Harmon. 2017. Estimating correlated rates of trait evolution with uncertainty. bioRxiv, 102939. OA

  • Duchen, P., C. Leuenberger, S. M. Szilágyi, L. J. Harmon, J. Eastman, M. Schweizer, and D. Wegmann. In press. Inference of evolutionary jumps in large phylogenies using Levy processes. Systematic Biology. OA.

  • Hagey, T. J., J. C. Uyeda, K. E. Crandell, J. A. Cheney, K. Autumn, and L. J. Harmon. In press. Tempo and mode of performance evolution across multiple independent origins of adhesive toe pads in lizards. Evolution. link - pdf.

  • Harmon, L. J. 2017. Contingent Predictability in Mammalian Evolution. Current Biology 27 (11), R425-R428. link - pdf.

  • Miller, E. T., D. N. Bonter, C. Eldermire, B. G. Freeman, E. I. Greig, L. J. Harmon, C. Lisle, and W. M. Hochachka. 2017. Fighting over food unites the birds of North America in a continental dominance hierarchy. bioRxiv, 104133. OA.

  • Miller, E. T., S. K. Wagner, L. J. Harmon, and R. E. Ricklefs. 2017. Radiating despite a Lack of Character: Ecological Divergence among Closely Related, Morphologically Similar Honeyeaters (Aves: Meliphagidae) Co-occurring in Arid Australian Environments. American Naturalist 189 (2), E14-E30. link - pdf.

  • Weber, M. G., C. E. Wagner, R. J. Best, L. J. Harmon, and B. Matthews. In press. Evolution in a community context: on integrating ecological interactions and macroevolution. Trends in Ecology & Evolution. link - pdf.


  • Des Roches, S., L. J. Harmon. and E. B. Rosenblum. 2016. Colonization of a novel depauperate habitat leads to trophic niche shifts in three desert lizard species. Oikos 125 (3), 343-353. link - pdf

  • Hagey, T. J., N. Cole, D. Davidson, A. Henricks, L. L. Harmon, and L. J. Harmon. 2016. Temporal Variation in Structural Microhabitat Use of Phelsuma Geckos in Mauritius. Journal of Herpetology 50 (1), 102-107. link - pdf.

  • Uyeda, J. C., L. J. Harmon, and C. E. Blank. A comprehensive study of cyanobacterial morphological and ecological evolutionary dynamics through deep geologic time. PloS ONE 11 (9), e0162539. OA


  • Deans, A. R. and 55 others. 2015. Finding our way through phenotypes. PLoS Biology 13: e1002033. OA

  • Des Roches, S., M. Brinkmeyer, L. J. Harmon, and E. B. Rosenblum. 2015. Ecological release and directional change in White Sands lizard trophic ecomorphology. Evolutionary Ecology 29: 1-16. link - pdf

  • Hardwick, K. M., L. J. Harmon, S. D. Hardwick, and E. B. Rosenblum. 2015. When field experiments yield unexpected results: lessons learned from measuring selection in White Sands lizards. PLoS ONE 10: e0118560. OA

  • Harmon, L. J. and S. Harrison. 2015. Species diversity is dynamic and unbounded at local and continental scales. The American Naturalist 185: 584-593. link - pdf - [counter argument] (

  • Nuismer, S. L. and L. J. Harmon. 2015. Predicting rates of interspecific interaction from phylogenetic trees. Ecology Letters 18: 17-28. link - pdf

  • Pennell, M.W. 2015. Modern Phylogenetic Comparative Methods and Their Application in Evolutionary Biology: Concepts and Practice.—Edited by László Zsolt Garamszegi (Book Review). Systematic Biology 64:161-163. link - pdf

  • Pennell, M. W., R. G. FitzJohn, W. K. Cornwell, and L. J. Harmon. 2015. Model adequacy and the macroevolution of angiosperm functional traits. American Naturalist 186 (2), E33-E50. preprint - github - poster

  • Pennell, M.W.*, M. Kirkpatrick*, S.P. Otto*, J.C. Vamosi, C.L. Piechel, N. Valenzuela, and J. Kitano. 2015. Y fuse? Sex chromosome fusions in fishes and reptiles. PLoS Genetics 11:e1005237. link - pdf

  • Rosindell, J., L. J. Harmon, and R. S. Etienne. 2015. Unifying ecology and macroevolution with individual-based theory. Ecology Letters 18: 472-482. link - pdf

  • Tank, D. C., J. M. Eastman, M. W. Pennell, P. S. Soltis, D. E. Soltis, C. E. Hinchliff, J. W. Brown, and L. J. Harmon. In press. Nested radiations and the pulse of angiosperm diversification. New Phytologist. link - pdf

  • Uyeda, J.C., D.S. Caetano, and M.W. Pennell. In press. Comparative analysis of principal components can be misleading. Systematic Biology. link - pdf - github


  • Anderson, C. J. R. and L. J. Harmon. 2014. Ecological and mutation-order speciation in digital organisms. American Naturalist 183: 257-268. link - pdf

  • Cornwell, W.K., M. Westoby, D.S. Falster, R.G. FitzJohn, B.C. O’Meara, M.W. Pennell, D.J. McGlinn, J.M. Eastman, A.T. Moles, P.B. Reich, D.C. Tank, I.J. Wright, L.Aarssen, J.M. Beaulieu, R.M. Kooyman, M.R. Leishman, E.T. Miller, U. Niinemets, J. Oleksyn, A. Ordonez, D.L. Royer, S.A. Smith, P.F. Stevens, L. Warman, P. Wilf, and A.E. Zanne. 2014. Functional distinctiveness of major plant lineages. Journal of Ecology 102:345-356. link - pdf

  • Cranston, K., L. J. Harmon, M. A. O’Leary, and C. Lisle. 2014. Best practices for data sharing in phylogenetic research. PLoS Currents Tree of Life 2014 Jun 19. OA

  • Davis, C. C., H. Schaefer, W. R. Anderson, Z. Xi, D. A. Baum, M. J. Donoghue, and L J. Harmon. 2014. Long-term morphological stasis maintained by a plant-pollinator mutualism. PNAS 111: 5914-5919. link - pdf

  • DesRoches, S., J. Torresdal, T. Morgan, L. J. Harmon, and E. B. Rosenblum. 2014. Beyond black and white: divergent behavior and performance in three rapidly evolving lizard species at White Sands. Biological Journal of the Linnean Society 111: 169-182. link - pdf

  • FitzJohn, R.G.*, M.W. Pennell*, A.E. Zanne, P.F. Stevens, D.C. Tank, and W.K. Cornwell. 2014. How much of the world is woody? Journal of Ecology 102:1266-1272. link - pdf - github - site - blog1 - blog2

  • Hagey, T., J. Puthoff, M. Holbrook, L. J. Harmon, and K. Autumn. 2014. Variation in Setal Micromechanics and Performance of Two Gecko Species. Zoomorphology 133: 111-126. link - pdf

  • Jochimsen, D. M., C. R. Peterson, and L. J. Harmon. 2014. Influence of ecology and landscape on snake road mortality in a sagebrush-steppe ecosystem. Animal Conservation 17: 583-592. link - pdf

  • Lanfear, R. and M.W. Pennell. 2014. Open access is worth considering. Trends in Plant Sciences 19:340-341. link - pdf

  • Muschick, M., P. Nosil, M. Roesti, M. T. Dittman, L. Harmon, and W. Salzburger. 2014. Testing the stages model in the adaptive radiation of cichlid fishes in East African Lake Tanganyika. Proceedings of the Royal Society B 281: 20140605. link - pdf

  • Pennell, M. W., J. M. Eastman, G. J. Slater, J. W. Brown, J. C. Uyeda, R. G. Fitzjohn, M. E. Alfaro, and L. J. Harmon. 2014. geiger v2.0: an expanded suite of methods for fitting macroevolutionary models to phylogenetic trees. Bioinformatics 30: 2216-2218. link - pdf

  • Pennell, M. W., L. J. Harmon, and J. C. Uyeda. 2014. Is there room for punctuated equilibrium in macroevolution? Trends in Ecology and Evolution 29: 23-32. link - pdf - rebuttal 1 - rebuttal 2 - reply

  • Pennell, M. W., L. J. Harmon, and J. C. Uyeda. 2014. Speciation is unlikely to drive divergence rates. Trends in Ecology and Evolution 29: 72-73. link - pdf

  • Stansbury, C.R., D.E. Ausband, P. Zager, C.M. Mack, C.R. Miller, M.W. Pennell, and L.P. Waits. 2014. A long term population monitoring approach to a wide-ranging carnivore: noninvasive genetic sampling of gray wolf rendezvous sites in Idaho, U.S.A. Journal of Wildlife Management 78:1040-1049. link - pdf - data

  • The Tree of Sex Consortium; T. Ashman, D. Bachtrog, H. Blackmon, E.E. Goldberg, M.W. Hahn, M. Kirkpatrick, J. Kitano, J.E. Mank, I. Mayrose, R. Ming, S.P. Otto, C.L. Peichel, M.W. Pennell, N. Perrin, L. Ross, N. Valenzuela, and J.C. Vamosi. 2014. Tree of Sex: a database of sexual systems. Scientific Data 1:140015. link - pdf - website - database

  • Uyeda, J. C. and L. J. Harmon. 2014. A novel Bayesian method for inferring and interpreting the dynamics of adaptive landscapes from phylogenetic comparative data. Systematic Biology 63: 902-918. link - pdf

  • Wagner, C. E., L. J. Harmon, and O. Seehausen. 2014. Cichlid species-area curves are shaped by adaptive radiations that scale with area. Ecology Letters 17: 538-592. link - pdf


  • Des Roches, S., J. B. Shurin, D. Schluter, and L. J. Harmon. 2013. Ecological and evolutionary effects of stickleback on ecosystem function. PLoS ONE 8: e59644. OA

  • Eastman, J. M., L. J. Harmon, and D. C. Tank. 2013. Congruification: support for time-scaling large phylogenetic trees. Methods in Ecology and Evolution 4: 688-691. link - pdf

  • Hardwick, K.M., J.M. Robertson, and E.B. Rosenblum. 2013. Asymmetrical mate preference in recently adapted white sands and black lava populations of Sceloporus undulatus. Current Zoology, Special Issue on Ecological Speciation, 59: 20-30. link - pdf

  • Harmon, L. J. 2013. Is character displacement central to ecology and evolutionary biology? Ecology 94: 2658-2659. link - pdf

  • Harmon, L. J. 2013. “Convergent Evolution.” In Oxford Bibliographies in Evolutionary Biology. Ed. Jonathan Losos. New York: Oxford University Press. link - pdf

  • Harmon, L. J. 2013. Macroevolutionary rates. In: J. B. Losos, Ed., The Princeton Guide to Evolution. Princeton University Press. link to book -

  • Harmon, L. J., J. Baumes, C. Hughes, J. Soberon, C. Specht, B. Thacker, W. Turner, and C. Lisle. 2013. Arbor: Comparative Analysis Workflows for the Tree of Life. PLoS Currents: Tree of Life: June 21, 2013. OA

  • Maliska, M.E., M.W. Pennell, and B.J. Swalla. 2013. Developmental mode influences diversification in ascidians. Biology Letters 9:20130068. link - pdf

  • Pennell, M. W. and L. J. Harmon. 2013. An Integrative View of Phylogenetic Comparative Methods: Connections to Population Genetics, Paleobiology and Community Ecology. The Year in Evolutionary Biology 1289: 90-105. link - pdf

  • Rosindell, J. and L. J. Harmon. 2013. A unified model of species immigration, extinction and abundance on islands. Journal of Biogeography 40: 1107-118. link - pdf

  • Slater, G. J. and L. J. Harmon. 2013. Unifying fossils and phylogenies for comparative analyses of diversification and trait evolution. Methods in Ecology and Evolution 4: 699-702. link - pdf

  • Stoltzfus, A., H. Lapp, N. Matasci, H. Deus, B. Sidlauskas, C.M. Zmasek, G. Vaidya, E. Pontelli, K. Cranston, R. Vos, C. O. Webb, L. J. Harmon, M. Pirrung, B. O’Meara, M. W. Pennell, S. Mirarab, M. S. Rosenberg, J. P. Balhoff, H. M. Bik, T. Heath, P. Midford, J. W. Brown, E. J. McTavish, J. Sukumaran, M. Westneat, M. E. Alfaro, and A. Steele. 2013. Phylotastic! Making Tree-of-Life Knowledge Accessible, Reusable and Convenient. BMC Bioinformatics 14:158. OA

  • Tyerman, J. G., J. M. Ponciano, P. Joyce, L. J. Forney and L. J. Harmon. 2013. The evolution of antibiotic susceptibility and resistance during the formation of Escherichia coli biofilms. BMC Evolutionary Biology 13: 22. OA


  • Godsoe, W. and L. J. Harmon. 2012. How do species interactions affect species distribution models? Ecography 35: 811-820. link - pdf

  • Harmon, L. J. 2012. An inordinate fondness for eukaryotic diversity. PLoS Biology 10: e1001382. OA

  • Ingram. T., L. J. Harmon, and J. B. Shurin. 2012. When should we expect early bursts of trait evolution in comparative data? Predictions from an evolutionary food web model. Journal of Evolutionary Biology 25: 1902-1910. link - pdf

  • Pennell, M.W. 2012. Biology in the light of phylogeny. Trends in Ecology & Evolution 27:657-658. link - pdf

  • Pennell, M. W., B. A. J. Sarver, and L. J. Harmon. 2012. Trees of Unusual Size: Sampling Bias Can Influence Inference of Early Bursts from Molecular Phylogenies. PLoS ONE 7: e43348. OA

  • Pennell, M.W., C.R. Stansbury, L.P. Waits, and C.R. Miller. 2012. Capwire: a R package for estimating population census size from non-invasive genetic sampling. Molecular Ecology Resources 13:154-157. link - pdf

  • Rosenblum, E. B., B. A. J. Sarver, J. W. Brown, S. Des Roches, K. M. Hardwick, T. D. Hether, J. M. Eastman, M. W. Pennell, and L. J. Harmon. 2012. Goldilocks meets Santa Rosalia: An ephemeral speciation model explains patterns of diversification across time scales. Evolutionary Biology 39: 255-261 OA

  • Rosindell, J., S. P. Hubbell, F. He, L. J. Harmon, and R. S. Etienne. 2012. The case for ecological neutral theory. Trends in Ecology and Evolution 27: 203-208. link - pdf

  • Rosindell, J. and L. J. Harmon. 2012. OneZoom: A Fractal Explorer for the Tree of Life. PLoS Biology 10: e1001406. OA - OneZoom

  • Slater, G. J., L. J. Harmon, P. Joyce, L J. Revell, and M. E. Alfaro. 2012. Fitting models of continuous trait evolution to incompletely sampled comparative data using Approximate Bayesian Computation. Evolution 66: 752-762. link - pdf

  • Slater, G. J., L. J. Harmon, and M. E. Alfaro. 2012. Integrating Fossils with Molecular Phylogenies Improves Inference of Trait Evolution. Evolution 12: 3931-3944. link - pdf

  • Wagner, C. E., L. J. Harmon, and O. Seehausen. 2012. Ecological opportunity and sexual selection together predict adaptive radiation. Nature 487: 366-369. link - pdf


  • Akçay, C., W. Searcy, S.E. Campbell, V.A. Reed, C.N. Templeton, K.M. Hardwick, and M. Beecher. 2011. Who initiates extra-pair copulations in song sparrows? Behavioral Ecology 23: 44-50. link - pdf

  • Brock, C. D. , L. J. Harmon, and M. E. Alfaro. 2011. Testing for Temporal Variation in Diversification Rates When Sampling is Incomplete and Nonrandom. Systematic Biology 60: 410-419. link - pdf

  • Carlson, B. A., S. M. Hasan, M. Hollmann, D. B. Miller, L. J. Harmon, and M. E. Arnegard. 2011. Brain evolution triggers explosive diversification of species and signals. Science 332: 583-586. link - pdf

  • Davies, T. J., G. Smith, D. U. Bellstedt, J. Boatwright, B. Bytebier, R. Cowling, F. Forest, L. J. Harmon, A. M. Muasya, B. D. Schrire, Y. Steenkamp, M. van de Bank, and V. Savolainen. 2011. Extinction risk and diversification are linked in a plant biodiversity hotspot. PLoS Biology 9: e1000620. OA

  • Des Roches, S., J. M. Robertson, L. J. Harmon, and E. B. Rosenblum. 2011. Ecological release in white sands lizards. Ecology and Evolution 1: 571-578. OA

  • Eastman, J. M., M. E. Alfaro, P. Joyce, A. L. Hipp, and L. J. Harmon. 2011. A novel comparative method for modeling shifts in the rate of character evolution on trees. Evolution 65: 3578-3589. link - pdf

  • Eastman, J. M., L. J. Harmon, H.-J. La, P. Joyce, and L. J. Forney. 2011. The onion model, a simple neutral model for the evolution of diversity in bacterial biofilms. J. Evol. Biol. 11: 2496-2504. link - pdf

  • Harmon, L. J. 2011. My island life. In: J. B. Losos, editor. Evolution Emerging: Essays from Leading Evolutionary Biologists. Roberts & Company Publishers. book

  • McInnes, L., W. J. Baker, T. G. Barraclough, K. K. Dasmahapatra, A. Goswami, L. J. Harmon, H. Morlon, A. Purvis, J. Rosindell, G. H. Thomas, S. T. Turvey, and A. B. Phillimore. 2011. Integrating ecology into macroevolutionary research (meeting report). Biology Letters 7: 644-646. OA - all talks from the meeting - mine

  • Rosenblum, E. B. and L. J. Harmon. 2011. Same same but different: replicated ecological speciation at White Sands. Evolution 65: 946-960. link - pdf

  • Smith, K. L., L. J. Harmon, L. Shoo, and J. Melville. 2011. Evidence of constrained phenotypic evolution in a cryptic species complex of agamid lizards. Evolution 65: 976-992. link - pdf

  • Stack, J., L. J. Harmon, and B. O’Meara. 2011. RBrownie: An R package for testing hypotheses about rates of evolutionary change. Methods in Ecology and Evolution 2: 660-662. link - pdf

  • Harmon, L. J. 2011. My island life. In: J. B. Losos, editor. Evolution Emerging: Essays from Leading Evolutionary Biologists. Roberts & Company Publishers. book


  • Arnegard, M. E., P. B. McIntyre, L. J. Harmon, M. L. Zelditch, W. G. R. Crampton, J. K. Davis, J. P. Sullivan, S. Lavoué, and C. D. Hopkins. 2010. Sexual signal evolution outpaces ecological divergence during electric fish species radiation. American Naturalist 176:335-356. link - pdf

  • Hagey, T. J., J. B. Losos, and L. J. Harmon. 2010. Cruise foraging of invasive chameleons (Chamaeleo jacksonii) in Hawaii. Breviora 519:1-7. link - pdf

  • Harmon, L. J. 2010. Journal Club: An evolutionary biologist ponders the pace of evolution. Nature 463: 1003. link - pdf

  • Harmon, L. J. and S. Braude. 2010. Conservation of Small Populations: Effective Population Size, Inbreeding, and the 50/500 Rule. pp. 125-138 in S. Braude and B. S. Low, Eds., An Introduction to Methods and Models in Ecology and Conservation Biology. Princeton University Press. book

  • Harmon, L. J. and R. E. Glor. 2010. Poor statistical performance of the Mantel test in phylogenetic comparative analyses. Evolution 64: 2173-2178, doi:10.1111/j.1558-5646.2010.00973.x. link - pdf

  • Harmon, L. J., J. B. Losos, J. Davies, R. G. Gillespie, J. L. Gittleman, W. B. Jennings, K. Kozak, M. A. McPeek, F. Moreno-Roark, T. J. Near, A. Purvis, R. E. Ricklefs, D. Schluter, J. A. Schulte II, O. Seehausen, B. Sidlauskas, O. Torres-Carvajal, J. T. Weir, & A. Ø. Mooers. 2010. Early bursts of body size and shape evolution are rare in comparative data. Evolution 64: 2385-2396. doi:10.1111/j.1558-5646.2010.01025.x. link - pdf

  • Matthews, B., L. J. Harmon, L. M’Gonigle, K. B. Marchinko, and H. Schaschl. 2010. Sympatric and allopatric divergence of MHC genes in threespine stickleback. PLoS ONE 5:e10948. OA -

  • Yoder, J. B., S. Des Roches, J. M. Eastman, L. Gentry, W. K. W. Godsoe, T. Hagey, D. Jochimsen, B. P. Oswald, J. Robertson, B. A. J. Sarver, J. J. Schenk, S. F. Spear, and L. J. Harmon. 2010. Ecological opportunity and the origin of adaptive radiations. Journal of Evolutionary Biology 23: 1581-1596, doi:10.1111/j.1420-9101.2010.02029.x. link - pdf


  • Alfaro, M. E., F. Santini, C. Brock, H. Alamillo, A. Dornburg, D. L. Rabosky, G. Carnevale, and L. J. Harmon. 2009. Nine exceptional radiations plus high turnover explain species diversity in jawed vertebrates. PNAS 106:13410-13414. link - pdf

  • Bailey, J. K., A. Hendry, M. Kennison, D. Post, E. Palkovacs, F. Pelletier, L. J. Harmon, and J. A. Schweitzer. 2009. From Genes to Ecosystems: An Emerging Synthesis of Eco-Evolutionary Dynamics. New Phytologist 184: 746–749. link - pdf

  • Harmon, L. J.*, B. Matthews*, S. Des Roches, J. Chase, J. Shurin, and D. Schluter. 2009. Evolutionary diversification in stickleback affects ecosystem functioning. Nature 458: 1167-1170. link - pdf

  • Ingram, T., L. J. Harmon, and J. B. Shurin. 2009. Niche evolution, trophic structure and species turnover in model food webs. American Naturalist 174: 56-67. link - pdf

  • Nosil, P. and L. J. Harmon. 2009. Niche dimensionality and ecological speciation. In: R. Butlin, J. Bridle, and D. Schluter, editors. Speciation and Patterns of Diversity. Cambridge University Press. book -

  • Nosil, P., L. J. Harmon, and O. Seehausen. 2009. Ecological explanations for (incomplete) speciation. Trends in Ecology and Evolution 24:145-156. link - pdf

  • Santini, F., L. J. Harmon, G. Carnevale, and M. E. Alfaro. 2009. Did genome duplication drive the origin of teleosts? A comparative study of diversification in ray-finned fishes. BMC Evlutionary Biology 9: 194. OA


  • Harmon, L. J., J. Melville, A. Larson, and J. B. Losos. 2008. The Role of Geography and Ecological Opportunity in the Diversification of Day Geckos (Phelsuma). Systematic Biology 57: 562-573. link - pdf

  • Harmon, L. J., J. Weir, C. Brock, R. E. Glor, and W. Challenger. 2008. GEIGER: Investigating evolutionary radiations. Bioinformatics 24:129-131. link - pdf

  • Pinto, G., D. L. Mahler, L. J. Harmon, and J. B. Losos. 2008. Testing the island effect in adaptive radiation: rates and patterns of morphological diversification in Caribbean and mainland Anolis lizards. Proceedings of the Royal Society B 275: 2749-2757. link - pdf

  • Revell, L. J. and L. J. Harmon. 2008. Testing quantitative genetic hypotheses about the evolutionary rate matrix for continuous characters. Evolutionary Ecology Research 10: 311-321. link - pdf

  • Revell, L. J., L. J. Harmon, and D. C. Collar. 2008. Phylogenetic signal, evolutionary process, and rate. Systematic Biology 57: 591-601. link - pdf


  • Harmon, L. J., L. L. Harmon, and C. G. Jones. 2007. Competition and community structure in diurnal arboreal geckos (genus Phelsuma) in the Indian Ocean. Oikos 116: 1863-1878, DOI: 10.1111/j.2007.0030-1299.15958.x. link - pdf

  • Mooers, A.O., Harmon, L.J., Wong, D.H.J., and S.B. Heard. 2007. Some models of phylogenetic tree shape. Pages 147-168 in New Mathematical Models for Evolution (O. Gascuel and M. Steel, eds.), Oxford University Press, Oxford. book

  • Nicholson, K. E., L. J. Harmon, and J. B. Losos. 2007. Evolution of Anolis lizard dewlap diversity. PLoS ONE 2(3): e274. doi:10.1371/journal.pone.0000274. OA

  • Revell, L. J., L. J. Harmon, R. B. Langerhans, and J. J. Kolbe. 2007. A phylogenetic approach to determining the importance of constraint on phenotypic evolution in the neotropical lizard Anolis cristatellus. Evolutionary Ecology Research 9: 261-282. link - pdf

  • Vellend, M., L. J. Harmon, J. L. Lockwood, M. M. Mayfield, A. R. Hughes, J. P. Wares, and D. F. Sax. 2007. Effects of exotic species on evolutionary diversification. Trends in Ecology and Evolution 22: 481-488. link - pdf


  • Harmon, L. J. and R. Gibson. 2006. Multivariate phenotypic evolution among island and mainland populations of the ornate day gecko, Phelsuma ornata. Evolution 60: 2622-2632. link - pdf

  • Melville, J., L. J. Harmon, and J. B. Losos. 2006. Intercontinental community convergence of ecology and morphology in desert lizards. Proceedings of the Royal Society Series B: Biological Sciences 273: 557-563. link - pdf


  • Harmon, L. J., K. Bauman, M. McCloud, J. Parks, S. Howell, and J. B. Losos. 2005. What the free-ranging animals do at the zoo: a study of the behavior and habitat use of opossums (Didelphis virginiana) on the grounds of the St. Louis Zoo. Zoo Biology 24: 197-213. link - pdf

  • Harmon, L. J., J. J. Kolbe, J. M. Cheverud, and J. B. Losos. 2005. Convergence and the multidimensional niche. Evolution 59: 409-421. link - pdf

  • Harmon, L. J. and J. B. Losos. 2005. The effect of intraspecific sample size on type I and type II error rates in comparative studies. Evolution 59:2705-2710. link - pdf

  • Kozak, K. H., A. Larson, R. M. Bonett, and L. J. Harmon. 2005. Phylogenetic analysis of ecomorphological divergence, species coexistence, and diversification rates in dusky salamanders (Plethodontidae, Desmognathinae). Evolution 59: 2000-2016. link - pdf

  • Revell, L. J., L. J. Harmon, and R. E. Glor, 2005. Under-parameterized model of sequence evolution leads to bias in the estimation of diversification rates from molecular phylogenies. Systematic Biology 54: 973-983. link - pdf

  • Weisrock, D. W., L. J. Harmon, and A. Larson. 2005. Resolving deep phylogenetic relationships in salamanders: analyses of mitochondrial and nuclear genomic data. Systematic Biology 54:758-777. link - pdf


  • Harmon, L. J., J. A. Schulte, J. B. Losos, and A. Larson. 2003. Tempo and mode of evolutionary radiation in iguanian lizards. Science 301: 961-964. link - pdf


  • Harmon, L. J. 2002. Some observations of the natural history of the prehensile-tailed skink, Corucia zebrata, in the Solomon Islands. Herpetological Review 33: 177-179. pdf


  • Harmon, L. J. 2000. A Translocation Strategy for Confiscated Pancake Tortoises. Chelonian Conservation and Biology 3(4):738-743.


  • Kolbe, J. J., L. J. Harmon, and D. A. Warner. 1999. New state record lengths and associated natural history notes for some Illinois snakes. Transactions of the Illinois State Academy of Science 92:133-135. pdf